In both these species the gills distinctly touch and grow on to the stem.
Gills with free nonreflected filaments.
The dorsal skeletal elements of the thorax and of the anterior six abdominal segments unite with the wing-cases to form a large respiratory chamber, containing five pairs of tracheal gills, with lateral slits for the inflow and a posterior orifice for the outflow of water.
The embryo passes through three stages - (I) still enclosed within the egg and living on its own yolk; (2) free, within the vitelline mass, which is directly swallowed by the mouth; (3) there is no more vitelline mass, but the embryo is possessed of long external gills, which serve for an exchange of nutritive fluid through the maternal uterus, these gills functioning in the same way as the chorionic villi of the mammalian egg.
Surface of gills smooth, gill-filaments all similar, with interlamellar junctions.