First toe represented by a minute rudiment of a metacarpal bone; the fourth by a metacarpal and two small phalanges without a claw, and not reaching the middle of the metacarpal of the third; fifth entirely absent.
Tail of moderate length, thick at the base and tapering towards the apex, clothed with short hair_ First hind toe (including the metacarpal bone) absent.
The first metacarpal is short and fuses throughout its length with the second.
Reduction and final loss of outer pair of digits (second and fifth), with coalescence of the metacarpal and metatarsal bones of the two middle digits to form a cannon-bone.
The primitive Artiodactyla thus probably had the typical number (44) of incisor, canine and molar teeth, brachyodont molars, conical odontoid process, four distinct toes on each foot, with metacarpal, metatarsal and all the tarsal bones distinct, and no frontal appendages.