This conclusion has shown that the Euthyneura do not represent an archaic form of Gastropoda, but are themselves derived from streptoneurous forms. The difference between the two sub-classes has been shown to be slight; certain of the more archaic Tectibranchia (Actaeon) and Pulmonata (Chilina) still have the visceral commissure long and not untwisted.
The Pulmonata are, like the other Euthyneura, hermaphrodite, with elaborately developed copulatory organs and accessory glands.
In aquatic Pulmonata the osphradium is retained.
In some Pulmonata (snails) the foot is extended at right angles to the visceral hump, which rises from it in the form of a coil as in Streptoneura; in others the visceral hump is not elevated, but is extended with the foot, and the shell is small or absent (slugs).
Pulmonata are widely distinguished from a small number of Streptoneura at one time associated with them on account of their mantle-chamber being converted, as in Pulmonata, into a lung, and the ctenidium or branchial plume aborted.
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