In aquatic Pulmonata the osphradium is retained.
This conclusion has shown that the Euthyneura do not represent an archaic form of Gastropoda, but are themselves derived from streptoneurous forms. The difference between the two sub-classes has been shown to be slight; certain of the more archaic Tectibranchia (Actaeon) and Pulmonata (Chilina) still have the visceral commissure long and not untwisted.
The genital duct is now said to be diaulic, as in Valvata, Oncidiopsis, Actaeon, and Lobiger among the Bullomorpha, in the Pleurobranchidae, in the Nudibranchia, except the Doridomorpha and most of the Elysiomorpha, and in the Pulmonata.
The Euthyneura comprises two orders, Opisthobranchia and Pulmonata.
An operculum is present only in Amphibola; a contrast being thus afforded with the operculate pulmonate Streptoneura (Cyclostoma, &c.), which differ in other essential features of structure from the Pulmonata.