In common with the okapi, giraffes have skin-covered horns on the head, but in these animals, which form the genus Giraffa, these appendages are present in both sexes; and there is often an unpaired one in advance of the pair on the forehead.
Uzel, with the majority of students, regards the paired organs as mandibles and the unpaired as an epipharynx.
Hinds believe that the paired piercers are the inner lobes of the maxillae, and the unpaired piercer the left mandible, the right mandible being absent.
C. Borner has stated that the unpaired piercer is attached directly to the base of the left maxilla.
The orbito-sphenoids diverge only posteriorly, otherwise they are practically unpaired and form the median interorbital septum, which is very large in correlation with the extraordinary size of the eyeballs.