Some members of this order spend the winter in the adult state, others in the " larval " or " pupal " condition.
After another moult the insect passes into the passive nymphal or " pupal " stage, during which it takes no food and rests in some safe hiding-place, such as the soil at the base of its food-plant or the hollow of a leaf-stalk.
In this it remains until the completion of the transformation into the sexually mature insect, which then emerges from the case, leaving the pupal integument behind.
As regards growth after hatching, all beetles undergo a "complete" metamorphosis, the wing-rudiments developing beneath the cuticle throughout the larval stages, and a resting pupal stage intervening between the last larval instal1 and the imago.
The pupal stage is passed in an earthen cell, just beneath the surface of the ground.