In origin the vascular system is due to a fusion of spaces which arise in the mesoblast of the larva.
Forming the mesoblast from ft, Foot.
Bergh (for Lumbricus and Criodrilus), whose figures show a derivation of the entire nephridium from mesoblast, and an absence of any connexion between successive nephridia by any continuous band, epiblastic or mesoblastic. A midway position is taken up by Wilson, who asserts the mesoblastic formation of the funnel, but also asserts the presence of a continuous band of epiblast from which certainly the terminal vesicle of the nephridium, and doubtfully the glandular part of the tube is derived.
The segmentation is of the mesoblast to begin with, and appears later behind the mouth, the part anterior to this becoming the prostomium of the adult.
The rhynchocoel is formed by a split which appears in the mesoblast surrounding the epiblastic pit which is the forerunner of the proboscis.