Flight-feather meaning

Any of the comparatively large, stiff feathers of a bird's wing or tail that are necessary for flight.
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Any of the large feathers of the wings or tail that support a bird in flight.
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Any of the comparatively large, stiff feathers of a bird's wing or tail that are necessary for flight.
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(ornithology) Any asymmetrical feather of the wing or tail of a bird.
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