This may possibly be the cell sap in their interior, which must exercise a slightly different hydrostatic pressure on the basal and, the lateral walls of the cells.
Schimper had previously maintained that the action of common salt in the cell-sap is detrimental as regards assimilation.
Young cells ar full of cytoplasm, old cells generally contain a large vacuole or vacuoles, containing cell-sap, and with only a thin, almost invisible layer of cytoplasm on their walls.
They are spherical, oval, fusiform, or rod-like, and are always found in the cytoplasm, never in the cell-sap. They appear to be permanent organs of the cell, and are transmitted from one cell to another by division.
The cell sap contains various substances in solution such as sugars, inulin, alkaloids, glucosides, organic acids and various inorganic salts.