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This in itself was not surprising, since plants normally fix CO2 in the dark using PEP carboxylase as part of their respiratory pathway.
The outer layer (the mesophyll) fixes CO2 using the enzyme PEP carboxylase to form 4-carbon organic acids.
This carbon-fixing reaction is catalyzed by the enzyme ribulose bisphosphate carboxylase, always known as rubisco.
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