Lameere's classification is founded on the number of abdominal sterna, the nervuration of the wings, the number of malpighian tubules (whether four or six) and other structural characters.
They may be distinguished from the Malacodermata by the presence of only five or six abdominal sterna, while six malpighian tubes are present in some of the families.
- This is an important tribe of beetles, including families with four malpighian tubes and only five or six abdominal sterna, while in the thorax there is a backwardly directed process of the prosternum that fits into a mesosternal cavity.
In the abdominal exoskeleton the segmental structure is very clearly marked, a series of sclerites - dorsal terga and abdominal sterna - being connected by pale, feebly chitinized cuticle, so that considerable freedom of movement between the segments is possible.
The first and second abdominal sterna are often suppressed or reduced, on account of the strong development of the hind-legs.