The ersaeome (Calycophorida), made up of the same appendages as the preceding type but with the addition of a nectocalyx; when free termed Ersaea.
The subsequent development is slightly different according as the future cormus is headed by a pneumatophore (Physophorida, Cystophorida) or by a nectocalyx (Calycophorida).
The planula develops, on the whole, in a similar manner, but the ectodermal invagination arises, not at the pole of the planula, but on the side of its broader portion, and gives rise, not to a pneumatophore, but to a nectocalyx, the primary swimming bell or protocodon (" Fallschirm ") which is later thrown off and replaced by secondary swimming bells, metacodons, budded from the coenosarc.
An opening to the exterior S From a comparison of the two embryological types there can be no doubt on two points; first, that the pneumatophore and the protocodon are strictly homologous, and, therefore if the nectocalyx is comparable to the umbrella of a medusa, as seems obvious, the pneumatophore must be so too; secondly, that the coenosarcal axis arises from the ex-umbrella of the medusa and cannot be compared to a manubrium, but is strictly comparable to the " bud-spike " of a Narcomedusan.
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