missing link

something necessary for completing a series; specif., an animal not yet found in the fossil record, that is an intermediate form of two animals, as such an animal coming between an ape and a human

See missing link in American Heritage Dictionary 4

noun
  1. A theoretical primate postulated to bridge the evolutionary gap between the anthropoid apes and humans. Not in scientific use.
  2. Something lacking that is needed to complete a series.

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