In Ephedra helvetica, as described by Jaccard, no proembryo or suspensor is formed; but the most vigorous fertilized egg, after undergoing several divisions, becomes attached to a tissue, termed the columella, which serves the purpose of a primary suspensor; the columella appears to be formed by the lignification of certain cells in the central region of the embryo-sac. At a later stage some of the cells in the upper (micropylar) end of the embryo divide and undergo considerable elongation, serving the purpose of a secondary suspensor.
As in other cell-walls, so here the older membranes may be altered by deposits of various substances, such as resin, calcium oxalate, colouring matters; or more profoundly altered throughout, or in definite layers, by lignification, suberization (Trametes, Daedalea), or swelling to a gelatinous mucilage (Tremella, Gymnosporangium), while cutinization of the outer layers is common.
The cell-membrane may become modified by the process of lignification, suberization, cuticularization or gelatinization.
Similarly, the small amount of cuticular and of epidermal protection, and of lignification in succulent halophytes may also be related to the same circumstance.
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