The latter has established, for all the Palaeozoic insects, an order Palaeodictyoptera, there being a closer similarity between the fore-wings and the hind-wings than is to be seen in most living orders of Hexapoda, while affinities are shown to several of these orders - notably the Orthoptera, Ephemeroptera, Odonata and Hemiptera.
In the Jurassic rocks are found Ephemeroptera and Odonata, as well as Hemiptera, referable to existing families, some representatives of which had already appeared in.
The sub-imago of the Ephemeroptera suggests that a moult, after the wings had become functional, was at one time general among the Hexapoda, and that the resting nymph of the Thysanoptera or the pupa of the Endopterygota represents a formerly active stage in the life-history.
As shown by the number and variety of species, the Orthoptera are the most dominant order of this group. Eminently terrestrial in habit, the differentiation of their fore-wings and hindwings can be traced from Carboniferous, isopteroid ancestors through intermediate Mesozoic forms. The Plecoptera resemble the Ephemeroptera and Odonata in the aquatic habits of their larvae, and by the occasional presence of tufted thoracic gills in the imago exhibit an aquatic character unknown in any other winged insects.
The Odonata are in many imaginal and larval characters highly specialized; yet they probably arose with the Ephemeroptera as a divergent offshoot of the same primitive isopteroid stock which developed more directly into the living Isoptera, Plecoptera, Dermaptera and Orthoptera.