The anatomical construction of these plants presents many peculiarities which have given rise to discussion as to the allocation of the order among the dicotyledons or among the monocotyledons, the general balance of opinion being in favour of the former view.
The typical structure of the vascular cylinder of the adult primary stem in the Gyrnnosperms and Dicotyledons is, like that of the higher ferns, a hollow cylinder of vas- Structure of cular tissue enclosing a central parenchymatous pith.
The outermost is the caiyptrogen, which gives rise to the root-cap, and in Dicotyledons to the piliferous layer as well.
In most of the existing Pteridophytes, in the Monocotyledons and in annual plants among the Dicotyledons, there is n further growth of much structural importance in the ~ d ~
Stems. But in nearly all perennial Dicotyledons, in all dicotyledonous and gymnospermous trees and shrubs and in fossil Pteridophytes belonging to all the great groups, certain layers of cells remain meristematic among the permanent tissues, or after passing through a resting stage reacquire menstematic properties, and give rise to secondary tissues.