The invaginated cells (derived from the division of the four big cells) form the endoderm or arch-enteron; the outer cells are the ectoderm.
Ing gs, the bilobed arch sh, The primitive shell-sac or enteron or lateral vesicles shell-gland ° of invaginated endoderm, pi, The rectal peduncle or whichwill develop into liver.
The other extreme end closes, but the invaginated endoderm cells remain in continuity with this extremity of the blastopore, and form the " rectal peduncle " or " pedicle of invagination " of Lankester, although the endoderm cells retain no contact with the middle region of the now closed-up blastopore.
The body-cavity and the muscular, fibrous and vascular tissues are traced partly to two symmetrically disposed " mesoblasts," which bud off from the invaginated arch-enteron, partly to cells derived from the ectoderm, which at a very early stage is connected by long processes with the invaginated endoderm.
The mass of the arch-enteron or invaginated endodermal sac has taken on a bilobed form, and its cells are swollen (gs and tge).