It is possible, however, that the absence of sunken stomata, and the occurrence of some other halophytic features, are related merely to the succulent habit and not to halophytism, for succulent species often occur on non-saline soils.
Warming states that the stomata of true, succulent, littoral halophytic herbs, in cases so far investigated, are not sunken (1909: 221).
Warming (1909: 220) quotes Griffon (1898), to the effect that the assimilatory activity is less in the halophytic form than in the ordinary form of the same species.