It appears that the Malpighian tubes of Scorpio are developed from the mesenteron, viz.
They have appeared independently in connexion with a change in the excretion of nitrogenous waste in Arachnids, Crustacea, and the other classes of Arthropoda when aerial, as opposed to aquatic, respiration has been established - and they have been formed in some cases from the mesenteron, in other cases from the proctodaeum.
The hypoblast is formed either by a definite invagination or by the immigration of isolated cells, known as vitellophags, which wander through the yolk and later become associated into a definite mesenteron, or by some combination of these two methods.
Renal excretory caeca (Malpighian tubes) are developed from the proctodaeum (not from mesenteron as in scorpion and Amphipoda).
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