If the three principal organ-systems of the medusa, namely mouth, tentacles and umbrella, be considered in the light of phylogeny, it is evident that the manubrium bearing the mouth must be the oldest, as representing a common property of all the Coelentera, even of the gastrula embryo of all Enterozoa.
The special property of the medusa is the umbrella, distinguishing the medusa at once from other morphological types among the Coelentera.
Some amongst them (Tergipes, Eolis) are also remarkable for possessing peculiarly modified cells placed in sacs (cnidosacs) at the apices of these same papillae, which resemble the " thread-cells " of the Coelentera.
Polypi (including the Coelentera of later authorities and the Polyzoa).
Of these latter, two grades were further distinguished by Lankester - those which remain possessed of a single archenteric cavity and of two primary cell-layers (the Coelentera or Diploblastica), and those which by nipping off the archenteron give rise to two cavities, the coelom or body-cavity and the metenteron or gut (Coelomata or Triploblastica).