The dorsal segments are sterile, corresponding to the bracts of Sphenophyllum Dawsoni, while the ventral segments constitute peltate sporangiophores, each bearing four sporangia, just as in a ax FIG.
Sp, Sporangiophores, each bearing a sporangium, sm.
The great length and slender proportions of the segments give the cone a peculiar character, but the relations of position appear to leave no doubt as to the homologies with the fructification of Sphenophylleae; as regards the sporangiophores, Bowmanites Romeri occupies exactly the middle place between S.
While the anatomy has a somewhat Lycopodiaceous character, the arrangement of the appendages is altogether that of the Sphenophylleae; at the same time Calamarian affinities are indicated by the characters of the sporangiophores and sporangia.
The sporangiophores, which are usually half as numerous in each verticil as the bracts, have the same form as in Equisetum, but each bears four sporangia only.