Rounded pear-shape, and when mature splits into two, exposing a crimson arillus surrounding a single seed (figs.
When the fruit is collected the pericarp is first removed; then the arillus is carefully stripped off and dried, in which state it forms the mace of commerce.
Ripens into a berry-like, very rarely capsular, fruit with the three groups of seeds arranged in lines along the walls, but imbedded in a pulpy arillus derived from the stalk of the seed.
The morphology of the female flowers has been variously interpreted by botanists; the peduncle bearing the ovules has been described as homologous with the petiole of a foliage-leaf and as a shoot-structure, the collar-like envelope at the base of the ovules being referred to as a second integument or arillus, or as the representative of a carpel.
The female flowers of the Taxaceae assume another form; in Microcachrys (Tasmania) the reproductive structures are spirally disposed, and form small globular cones made up of red fleshy scales, to each of which is attached a single ovule enclosed by an integument and partially invested by an arillus; in Dacrydium the carpellary leaves are very similar to the foliage leaves - each bears one ovule with two integuments, the outer of which constitutes an arillus.