A crop-like dilatation of the gut and a recurved intestine, embedded in the compact yellowish-brown liver, the ducts of which open into it, form the rest of the digestive tract and occupy a large bulk of the visceral hump. The buccal region presents a pair of shelly jaws placed laterally upon the lips, and a wide range of variation in the form of the denticles of the lingual ribbon or radula.
High and bears large pear-shaped fruits, green or deep purple in colour, with a firm yellowish-green marrow-like pulp surrounding a large seed.
The principal symptom may show itself in general pallor, including all cases where the normal healthy green hue is replaced by a sickly yellowish hue indicating that the chlorophyll apparatus is deficient.
The eggs deposited by the beetle in the ground develop into yellowish-brown wire-like grubs with six legs on the first three segments and a ventral prominence on the anal segment.