In the deer of the sambar group, where the antlers never advance beyond a three-tined type, the shedding is frequently, if not invariably, very irregular; but in the majority at least of the species with complex antlers the replacement is annual, the new appendages attaining their full development immediately before the pairing-season.
- Antlers smaller and simpler, four-tined, with a trez (third), but no bez (second); coat of adult spotted, at least in summer, with a white area bordered by black in the region of the tail, which is also black and white.
The rusine or sambar group of Cervus, of which the characteristics are given above, comprises a considerable number of longtailed species with three-tined antlers from the Indo-Malay countries and some parts of China.
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