In places the nematocysts may be crowded so thickly as to form a tough, supporting, " chondral " tissue, resembling cartilage, chiefly developed at the margin of the umbrella and forming streaks or bars supporting the tentacles (" Tentakelspangen," peronia) or the tentaculocysts (" Gehorspangen," otoporpae).
Brooks regards these organs as sensory, serving for the sense of balance, and representing a primitive stage of the tentaculocysts of Trachylinae; Linko, on the other hand, finding no nerve-elements connected with them, regards them as digestive (?) in function.
43, A); in this the mouth is formed distally as a perforation (B); next the sides of the tube so formed bulge out laterally near the attachment to form the umbrella, while the distal undilated portion of the tube represents the manubrium (C); the umbrella now grows out into a number of lobes or lappets, and the tentacles and tentaculocysts grow out, the former in a notch between two lappets, the latter on the apex of each lappet (D, E); finally, the velum arises as a growth of the ectoderm alone, the whole bud shapes itself, so to speak, and the little medusa is separated off by rupture of the thin stalk connecting it with the parent (F).
Medusae with sense-organs represented by otocysts derived from modified tentacles (tentaculocysts), containing otoliths of endodermal origin, and innervated from the ex-umbral nerve-ring.
Not lobed or indented; tentaculocysts usually enclosed in vesicles; gonads on the radial canals.