The character of this skull and the compound rhamphotheca (known by the imprints left upon the jaws) indicate affinities with the Steganopodes.
Rhamphotheca compound; cosmopolitan.
A diagnosis covering all the Ratitae (struthio, rhea, casuarius, dromaeus, apteryx and the allied fossils dinornis and aepyornis) would be as follows - (i) terrestrial birds without keel to the sternum, absolutely flightless; (ii) quadrate bone with a single proximal articulating knob; (iii) coracoid and scapula fused together and forming an open angle; (iv) normally without a pygostyle; (v) with an incisura ischiadica; (vi) rhamphotheca compound; (vii) without apteria or bare spaces in the plumage; (viii) with a complete copulatory organ, moved by skeletal muscles.
Skull bird-like, except that the short bill cannot have been enclosed in a horny rhamphotheca, since the upper jaw shows a row of 13, the lower jaw 3 conical teeth, all implanted in distinct sockets.
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