It has, however, been alleged that this cavity is formed by a pair of mesoblastic somites (N.
Bergh (for Lumbricus and Criodrilus), whose figures show a derivation of the entire nephridium from mesoblast, and an absence of any connexion between successive nephridia by any continuous band, epiblastic or mesoblastic. A midway position is taken up by Wilson, who asserts the mesoblastic formation of the funnel, but also asserts the presence of a continuous band of epiblast from which certainly the terminal vesicle of the nephridium, and doubtfully the glandular part of the tube is derived.
We have thus the replacement of a spermatheca, corresponding to those of the remaining families of Oligochaeta, and derived, as is believed, from the epidermis, by a structure performing the same function, but derived from the mesoblastic tissues, and with a cavity which is coelom.
Re, Stiebel's canal (left side), tge, Mesoblastic (skeletotrophic probably an evanescent and muscular) cells invest embryonic nephridium.
In the direct development Bateson showed that the three divisions of the coelom arise as pouches constricted off from the archenteron or primitive gut, thus resembling the development of the mesoblastic somites of Amphioxus.