2, nch.), growing forwards in the septum between the proboscis-cavity and the collar-cavities, and supported dorsally by the median mesentery of the collar, is the representative of the so-called notochord or stomochord of Balanoglossus; and if the view that this organ is really a notochord is well founded, it may be regarded as the homologue of the anterior end of the Vertebrate notochord.
It receives the three great venous trunks of the body, namely the vena cava superior dextra, the vena cava superior sinistra more dorsally, and the vena cava inferior more to the right and below; the opening of the last is guarded by two prominent valves in place of the mammalian valvula Eustachii.
Commonly the nephridia are strictly paired a single pair to each segment, while the branches of the blood vascular system are similarly metameric. The alimentary canal is nearly always a straight tube running from the mouth, which is surrounded by the first segment of the body and overhung by the prostomium, to the anus,which is then either surrounded by the last segment of the body or opens dorsally a little way in front of this.
Quite independent of these are certain large dorsally situate funnel-like folds of the coelomic epithelium, ciliated, but of which no duct has been discovered leading to the exterior.
The detorted visceral commissure shows a tendency to the concentration of all its elements round the oesophagus, so that except in the Bullomorpha and in Aplysia the whole nervous system is aggregated in the cephalic region, either dorsally or ventrally.