In ordinary language the name is used for any species of Siphonaptera (otherwise known as Aphaniptera), which, though formerly regarded as a suborder of Diptera, are now considered to be a separate order of insects.
The relation of the Diptera (two-winged flies, or flies proper) to the other Orders is dealt with underHexapoda.
The chief characteristic of the Diptera is expressed in the name of the Order, since, with the exception of certain aberrant and apterous forms, flies possess but a single pair of membranous wings, which are attached to the meso-thorax.
The wings, which are not capable of being folded, are usually transparent, but occasionally pigmented and adorned with coloured spots, blotches or bands; the wing-membrane, though sometimes clothed with minute hairs, seldom bears scales; the wing-veins, which are of great importance in the classification of Diptera, are usually few in number and chiefly longitudinal, there being a marked paucity of cross-veins.
The antennae of Diptera, which are also extremely important in classification, are thread-like in the more primitive families, such as the Tipulidae (daddy-long-legs), where they consist of a considerable number of joints, all of which except the first two, and sometimes also the last two, are similar in shape; in the more specialized families, such as the Tabanidae (horse-flies), Syrphidae (hover-flies) or Muscidae (house-flies, blue-bottles and their allies), the number of antennal joints is greatly reduced by coalescence, so that the antennae appear to consist of only three joints.