If the three principal organ-systems of the medusa, namely mouth, tentacles and umbrella, be considered in the light of phylogeny, it is evident that the manubrium bearing the mouth must be the oldest, as representing a common property of all the Coelentera, even of the gastrula embryo of all Enterozoa.
Polypi (including the Coelentera of later authorities and the Polyzoa).
Of these latter, two grades were further distinguished by Lankester - those which remain possessed of a single archenteric cavity and of two primary cell-layers (the Coelentera or Diploblastica), and those which by nipping off the archenteron give rise to two cavities, the coelom or body-cavity and the metenteron or gut (Coelomata or Triploblastica).
The Metazoa form two main branches; one, Parazoa, is but a small unproductive stock comprising only the Phylum Porifera or Sponges; the other, the great stem of the animal series Enterozoa, gives rise to a large number of diverging Phyla which it is necessary to assign to two levels or grades - a lower, Enterocoela (often called Coelentera), and a higher, Coelomocoela (often called Coelomata).
The structural features which the Mollusca do possess in common with other animals belonging to other great phyla of the animal kingdom are those characteristic of the Coelomata, one of the two great grades (the other and lower being that of the Coelentera) into which the higher animals; or Metazoa as distinguished from the Protozoa, are divided.
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