The barnacles and their allies, forming the group Cirripedia or Thyrostraca, sometimes treated as a separate sub-class, are distinguished by being sessile in the adult state, the larval antennules serving as organs of attachment, and the antennae being lost.
Eyestalks and the antennules in this aberrant group correspond to the primitive head somites or not, their distinctness is certainly a secondarily acquired character, for it is not found in the larvae, nor in any of the more primitive groups of Malacostraca.
This modification is usually only found in the antennules and antennae, but it may exceptionally be found in the appendages of the trunk, as, for instance, in the thoracic legs of some Decapods (e.g.
The antennules (or first antennae) are almost universally regarded as true appendages, though they differ from all the other appendages in the fact that they are always innervated from the " brain " (or preoral ganglia), and that they are uniramous in the nauplius larva and in all the Entomostracan orders.
As regards their innervation an apparent exception is found in the case of A pus, where the nerves to the antennules arise, behind the brain, from the oesophageal commissures, but this is, no doubt, a secondary condition, and the nerve-fibres have been traced forwards to centres within the brain.