The terminal phalanges of the four outer digits are small, somewhat conical and flattened in form.
The hind-limb is typically avine, with intertarsal joint, distally reduced fibula, and the three elongated metatarsals which show already considerable anchylosis; reduction of the toes to four, with 2, 3, 4 and 5 phalanges; the hallux is separate, and as usual in recent birds posterior in position.
First toe represented by a minute rudiment of a metacarpal bone; the fourth by a metacarpal and two small phalanges without a claw, and not reaching the middle of the metacarpal of the third; fifth entirely absent.
The first phalanges are expanded at their lower ends, and the wide, depressed middle phalanges embedded in a broad cutaneous pad, forming the sole of the foot, on which the animal rests in walking instead of on the hoofs.
The terminal phalanges are small and nodular, not flattened on their inner or opposed surfaces, and not completely encased in hoofs, but bearing nails on their upper surface only.
